Answer:
the correct answer is E
Explanation:
Althoug the other options may be correct, they would be indirect consequencies of the permeable membrane described.
The protons-gradient loss would affect immediately the ATP synthase. This enzymatic complex involves several proteins that act together: some of them change their three-dimentional conformation when protons pass through them and after this others are capable of catalyze the chemical bound between ADP and P (phosphate).